John is an Adjunct Professor of Physiology & Pharmacology at the Cumming School of Medicine at the University of Calgary. He holds a PhD in Medical Sciences (University of Toronto), a B.Sc. and an M.Sc. in Biological Sciences (Queen’s University). From 2009 to 2012, John was Founding Director of the Farncombe Family Digestive Health Research Institute at McMaster University. He completed post-doctoral studies at the Wellcome Research Laboratories in London, England, and he holds an MBA from the University of Birmingham, England.